Taxonomic Notes: Hordeum marinum Huds. is a tertiary wild relative of Barley, H. vulgare L. (USDA, ARS, National Genetic Resources Program ). Differs from Hordeum leporinum and Hordeum glaucum in that the junction of the with line drawing: Background and Aims Hordeum marinum. is a species complex that includes the diploid subspecies marinum and both diploid and tetraploid.

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Marnum 5 revealed the highest chromosomal variation both within and between taxa. If diploid gussoneanum and marinum were involved in the origin of tetraploid gussoneanumtheir chromosomes should be present in the latter.

A survey with and internationally agreed recommendation for barley chromosome nomenclature. Karyotype analysis of diploid gussoneanum Chromosome identification and karyotype analysis of two diploid accessions of gussoneanum BCC and GRA was performed by combining the physical patterns obtained with the same probes used to characterize marinum Fig.

However, the most submetacentric chromosome pair of gussoneanum also carried an interstitial site on the long arm Fig. In addition to the three 5SrDNA loci observed in the two pairs of satellitized chromosomes, hrodeum minor 5SrDNA locus was observed in a distal position on the long arm of a very metacentric chromosome Fig.

However, taking into account the diversity of karyotypes detected within marinumit cannot be ruled out that the non- gussoneanum genome will be found in other marinum ecotypes. An ecogeographical study of the genus Hordeum. Ecology An annual of barish places by the sea, on the trampled margins of dried-up pools and ditches in grazing marshes, on tracks and sea walls, and in the uppermost parts of saltmarshes; also, very locally, beside salt-treated roads inland.

Petersen G, Seberg O. Characteristic associates in these open communities include Puccinellia distans, Spergularia marina hordeumm the scarce species Polypogon monspeliensis, Puccinellia fasciculata and P. Progress in phylogenetic analysis and a new infrageneric classification of the barley genus Hordeum Poaceae: As maginum marinumAAC 5 produced a rich hybridization pattern of in situ signals.

Two 5SrDNA loci were located, one on the satellite of the satellitized chromosomes and one distal on the long arm of the smallest metacentric chromosome pair; i. Indeed, on the basis of molecular evidence, different authors have suggested the existence of two geographically separated marinum forms: New Hordeum – Triticum hybrids. The identification of each chromosome pair in all subspecies and cytotypes is reported for the first time.


Another major disagreement concerns the auto- or allopolyploid origin of the tetraploids.

Homologies were established among chromosomes of the different accessions. Their relationships, the rank of the taxa and the origin of the polyploid forms remain points of debate. In the present study, although polymorphic signals were detected between accessions BCC and GRA from Greece and Italy, respectively, the physical maps for each probe were very similar, but significantly different from that recorded for the Spanish accession GRA compare karyotypes in Fig.


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Perennation – primary Annual. Cytogenetic and Genome Research. To understand the genomic constitution of H. On the origin of the tetraploid species Hordeum capense and H. Species relationships between antifungal chitinase and nuclear rDNA internal transcribed spacer sequences in the genus Hordeum.

In conclusion, this study reveals the origin of the polyploid forms of H. Identification of the entire chromosome complement of bread wheat by two-colour FISH.

Molecular phylogeny and reticulate origins of several American polyploid Hordeum species. Karyotypes were constructed designating chromosomes from 1 to 7 on the basis of similar chromosome morphology, size and the physical mapping of the probes in marinum karyotypes Fig.

Karyotypes of amrinum two 2 hirdeum gussoneanum accessions studied showing one chromosome of each of the seven homologous pairs. Chromosome identities are indicated in white lettering. Origin of polyploids in H. When is enough, enough in phylogenetics? Relationships in the genus Hordeum: This is thought to be due to the rebuilding of sea defences, infilling of pools and ditches, the wholesale conversion of coastal grazing marshes to arable land, and, locally, the cessation of grazing.

Hordeum marinum Calflora

Chromosome identification and karyotype analysis of two diploid accessions of gussoneanum BCC and GRA was performed by combining the physical patterns obtained with the same probes used to characterize marinum Fig.

Assuming a monophyletic origin for the diploid cytotypes of H. Marinun and widely naturalised outside its native range. These are usually recognized as a single species, H.


Hordeum marinum

Skip to main content. However, several SSRs were found to provide useful diagnostic probes for chromosomal identification. The chromosomal distribution revealed by pAs1 was coincident with that seen for diploid H. Phylogenetic analyses of the diploid species of Hordeum Poaceae and a revised classification of the genus. The homologous relationship between chromosome pairs 6 and 7 was easily established using the rDNA loci as landmarks.

The aim of the present work was to identify hotdeum markers for characterizing the karyotypes of a representative sample of Marinuj. It is gordeum distributed in the Mediterranean region, where it grows on both maritime sands and on disturbed ground away tram the coast, and extends along the coast of western Europe to reach its northern limit in the British Isles.

This indicates that diversity exists among the marinum accessions. This is supported by the observation of regular meiotic behaviour in hybrids between marinum and diploid forms of gussoneanum e.

Phylogeny of two tetraploid Hordeum species, H. This is the first report amrinum identify all H. A third locus with no counterpart in marinum was located in a subterminal position on the long arm of the satellitized chromosome pair Fig.

Hordeum marinum – Wikipedia

The distribution, organization and evolution of two abundant and widespread repetitive DNA sequences in the genus Hordeum. The structure, amount and chromosomal localization of defined repeated DNA sequences in species of the genus Secale. AAC 5 signals were found at multiple sites, especially in the pericentromeric regions of most chromosomes; again, this rendered individual chromosome marinuum difficult Fig.

The distribution pattern of the set of repetitive DNA sequences, plus chromosome morphology, helped establish putative homologies among chromosomes assigned the same number in the subspecies of H. The taxonomic treatment of marinum and gussoneanum as subspecies marijum generally accepted.